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Excessive kneading can break down gluten strands, making the dough tough and elastic, leading to a dense final product. This happens when the gluten network is developed too much and then starts to degrade.
A dough with too little water will be stiff and difficult to work with, leading to a dense crumb. This can make it feel tough even if it's not over-kneaded.
Flours with higher protein content, like bread flour, develop gluten more readily. Using too much bread flour or over-kneading it can easily lead to toughness.
If the dough doesn't rise sufficiently before baking, the gluten structure won't have enough time to relax and expand, resulting in a dense loaf.
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